What is the The Guardian crossword answer for the clue: Loud, ultimately; lady ululating, primarily?

Loud, ultimately; lady ululating, primarily?

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Numerical homophone of 43-Across
Course in which integration is taught, for short
Santorini sandwiches
Start to rally, maybe
Endor inhabitants in the "Star Wars" universe
Ivory Coast's main export
Tank tops expose them
Miss ___ USA (beauty pageant)
Spot to swing from
Gives a shot
Exploratory trip
The "p" in Paddington Station?
Question that's a humble response to a clothing compliment
They put runners out
Retina contents
Actress Collette with an Emmy and a Golden Globe
Only M.L.B. player to have 50 home runs and 50 stolen bases in a single season
Show with an illustrious "Five-Timers Club," in brief
Put a pilot back into action?
Like the eyes of some Raggedy Anns
"Go on ahead"
Tree goop
El ___ (Six Flags coaster with horns on the front)
Source of many a motto
X or XL offering, perhaps
Major for someone who can correctly distinguish between Manet and Monet
Avoiding coverage, in a way
Nickname for basketball's Mike Krzyzewski
"When you teach your ___, you teach your ___'s ___": the Talmud
Adverb that's a homophone of 54-Down
Rightmost column on the periodic table
Wyoming's ___ Pass
Mustang, for one
One meaning of the "+" in LGBTQ+, in brief
Like many higher-ups
Roughly 120, for Vatican City

Loud, ultimately; lady ululating, primarily?

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